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A general approach to modelling of ecosystem dynamic is considered. The special term "coenome" is proposed for basic dynamic element of biogeocoenoses. The coenome is an association of populations which have complementary properties (up to symbioses) or united by trophical links. Each succession stage is characterised by a major dominant coenome. The succession process is considered as a process of step-by-step changing dominating coenome. Sketch models of the coenome are considered. Thedoi:10.7148/2012-0143-0149 dblp:conf/ecms/Chernyshenko12 fatcat:2l6mmmqppfgdpnlqslh2dbxgxe