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Functional magnetic resonance imaging (fMRI) based on the blood oxygen level dependent (BOLD) contrast has gained a prominent position in neuroscience for imaging neuronal activity and studying effective brain connectivity under working state and functional connectivity at resting state. However, the fundamental questions in regards to fMRI technology: how the BOLD signal inferences the underlying microscopic neuronal activity and physiological changes and what is the ultimate specificity ofdoi:10.1038/s41598-017-02864-0 pmid:28578390 pmcid:PMC5457450 fatcat:xwmhi2wpcrctzofkuott5h74zu