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e c o l o g i c a l i n d i c a t o r s 9 s ( 2 0 0 9 ) s 1 7 -s 2 8 (J.A. Browder). a v a i l a b l e a t w w w . s c i e n c e d i r e c t . c o m journal homepage: www.elsevier.com/locate/ecolind 1470-160X/$ -see front matter . Published by Elsevier Ltd.doi:10.1016/j.ecolind.2008.10.007 fatcat:dav2oyj4nfgzzdxefacziykutq