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Classical models of plant-insect coevolution have emphasized that related species of insects frequently exploit closely related groups of plants (Ehrlich and Raven 1964; Benson et al. 1976 ). Implicit in this concept is a series of adaptations and counteradaptations by both the host and the parasite. Such models, although descriptive of broad patterns in evolutionary time, do not address the ecological factors affecting individual fitness nor the genetic capacity for change. The purpose of thisdoi:10.1086/284633 fatcat:pkx2e6kb5bgxvniw4eu7tckfam