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8 Heavy-metal soil contamination is one of the major abiotic stress factors that, by 9 negatively affecting plant growth and development, severely limit agricultural 10 productivity worldwide. Plants have evolved various tolerance and detoxification 11 strategies in order to cope with heavy-metal toxicity, while ensuring adequate supply 12 of essential micronutrients at the whole-plant as well as cellular levels. Genetic 13 studies in the model plant Arabidopsis thaliana have been instrumentaldoi:10.1007/978-1-4939-3356-3_16 pmid:26867625 fatcat:nyucetgp7rbh7o7wdn5g3t3opm