Encephalartos middelburgensis (Zamiaceae): a new species from the Transvaal

P.J. Robbertse, Suzelle van der Westhuizen, P. Vorster
1989 South African Journal of Botany  
Encephalartos middelburgensis Vorster et al. is described from the upper reaches of the Olifants River. It resembles E. graniticolus Vorster et al., E. eugene-maraisii Verdoorn , E. verrucosus Vorster et al., E. lehmannii Lehm., E. princeps RA Dyer, and to some extent E. cupidus R.A. Dyer by its stiff, pungent, glaucous fronds; but differs from them all by its combination of well-developed petioles and proximal leaflets reduced in size to entire prickles towards base of frond. Anatomically it
more » ... ows an affinity to E. graniticolus, E. eugene-maraisii, and E. cupidus on account of its isobilateral, amphistomal leaflets. Encephalartos middelburgensis Vorster et al. word beskryf vanaf die bolope van die Olifantsrivier. Dit stem ooreen met E. graniticolus Vorster et al., E. eugene-maraisii Verdoorn, E. verrucosus Vorster et al., E. lehmannii Lehm ., E. princeps RA Dyer, en tot 'n mate E. cupidus RA Dyer wat betref die stywe, stekelpuntige, blougrys blare; maar verskil van hulle almal deur die kombinasie van 'n goed-ontwikkelde petiolus en proksimale pinnae wat na die basis toe kleiner word om gaafrandige stekels te vorm. Anatomies toon dit ooreenkoms met E. graniticolus, E. eugene-maraisii, en E. cupidus op grond van sy isobilaterale, amfistomale pinnae.
doi:10.1016/s0254-6299(16)31240-6 fatcat:2mfqbcsekjblpn66a5thsisq24