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Nine of the physiological leaf spot mutants were described and investigated linkage relation with marker genes. All of them were governed by single recessive genes respectively and the causal genes were designated as a series of spl (spotted leaves) as spl" spl,......sf&. With eight of them except for sPZ~, the linkage groups were determined as follows: sj+-a linkage group corresponding to A type of trisomics which never corresponded to any of twelve linkage groups published by Nagao anddoi:10.5109/23671 fatcat:24agmew36zcazn33jznq7g7e2q