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Water shortage is a critical problem touching plant growth and yield in arid and semi-arid region. Photosynthesis is one of the key processes to be affected by water deficits, via decreased CO 2 diffusion to the chloroplast and metabolic constraints. Understanding the physiological basis of drought tolerance will be helpful in can play a major role in stabilizing crop performance under drought and in the protection of natural vegetation. The functional state of photosynthetic apparatus is afatcat:3mbtfqvlpjf3poalfnit4cufai