ῒ ῍ ΐ ῐ ̮ ῭ῦ̮῭Ό QῨῲ̳̱QQQQ̯ῖῡΌ̱ ῏ ̲ ῴ῝΅QQῲ̲ῶ̱ ῝΅̮Qῌ ῗῤΊῢQQΊῧ QῩῦQ̱ῡ΅̮ῌ Ώ̯ῦQ̰ῥ̲̰Ῑ'΅῍ ̳̯ῧ ̳̱QQQῥQQ῝΅QQ̮ῬQQῲ΅῏Ίῠ̰QQῡ ̮΅ DNA ῦ Q ῧῘ̮῞̮ῡ΅̯̰ῌ QQῳ ῥ DNA QQῦQ῏Q̱ῲ῞̮Ί̯'ῠῤῢ̮Q̰ ῥῩ̯̮΅῍ ̮QῥῧῙ̰Ὶ̰ῤ̳̱Qῡ̳ῦῠQ̳Q ̮Q̰'ῠ΅̯̰ῌ ῟̯Ί̯ DNA ῲ̯QQ̮῞ῥῌ QQQῲQQΊῠ̯ῦ̳̳ῲ̳̱QQQῥQQ῟Ὶ΅ῲ ῷῤ̯̰ῡ΅῍ ̰̮̳̯ῦQ̰ῧQῳ̯QΊ̮̮Q ῥῤ̮ῠ̰̳̱QQQῡῧQ῏Ῑ'ῠΊ̰̮̱ῡ ΅῍ Q̲ῑQῌ QQῤΰ̮QῦQQῦ̰QῡQQΊ̯ QῦQQῲQῢΊῠΊ̰̮̯̰ῌ ̳̱QQQQῦ ῦ ῳῥῤ̯ῠ΅῍ ῧQῤ̮QῥῧQ̳Qῌ QQῤΰQ Qῦῡ̮ῲQῙ῝΅̳Q̮Qῦ῝΅̯̰ῌ ῗ'Ῥῧ̲Q QῦQ῏Q̱̰QῙ῟΅῍ ῗῦΏ̯ῦQ̰ῲQῥQQ῝΅̯̰ῥῌ "ῧ ̰QQQῲῌ῭´'ῠ῞̮̯Qῦ̳̱QῦῦQῲῲ῝ ΅QQῲQQΊῠ̮̯῍ 2ῐ4 QQῢ̮̰QQ̲Qῧ ῏Q̯ῦQQῌ ΅ῧ 1

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... ῼῦQQῡQ῏QQῲ῎QῙ ῟΅ῗῢῲQῳῢΊ̯̯̰ῡ΅῍ ῗῦQ ῦ̯̰ῥQ Qῌ Q῾Ίῠ΅QQῲQΊῌ ̮QῥQ̯ῌ Q̮ῦ ̯ ῥQ̯ῗῤῡQQῳῤQQQQῲQΎΊῌ Q ̰̳̯Ίῲ̰QΊ̯̯ ῲQQΊῠ̰Ῥ̮ῗῢῥΐΊῡ ̰QῥQ̯ῠ̯̯ῖ'ῨῢQ̮ῠ΅῍ 1. ῝῟Ῑ῏̮̮ῐῖ῞῎ῌῚῗῑΊῘ QQ̮ 16S rDNA ̲QῦῧQῥQ̯ῠQῴῳῥQ QῙ'΅Ύ̮ῥῤ῭ῌ QQQῳῥQQῙ'̯῝̰ῠῦ QῨῲ̳̱QῡQ῏῝΅ῗῢ̮ῡ̮΅Ύ̮ῥῤ̯̯῍ ῗ 'ῧ 1980 ̲ῒQ̮Ῥ ῷ̱ ῩQῦQQῴ ῏῾ῦ Ὼ῏ ῾ Ὸ̱̱̲ῥ̳̮ῢῗ̮Ῡ̮῍ 2009 ̲ 6 Qῦ̰QῡῬ̯Ῑ'̯ 16S rDNA ̲Qῧ 920, 643 Q ῦ̲Qῢ̲῾ῳῤῚῡῡQΊῠ΅῍ ῝QῥQQῙ' ̯Q῍ῧQQ 7,000 ῍̰ΰ΅ῦῡῌ Qῦ̮Qῌ Q̰ῦ Qῦ̮Q̰̯̮ῙῳῬ̯Ῑ'ῠ΅ῗῢῥῤ΅῍ Eproject ῡQQ῭ῦ ῞̳ῲQQῳῥQQ῝΅QQ̮ῶ ´Ῑ'ῠ΅̮ῌ ῪQQQῥῧQ̰ῤQQ῍̮̰̯Ῐ Q῍ῦQ῝΅ῢ'ῠ΅῍ QQῥ 16S rDNA ̯ῖ ῡQQῦQQῲῲΊΎ̮ῢ῝΅ῢQ̰ῦQ῍ῥQ῭ῤ ̮Q̲QῲQ̯̯Q̮QQῙ'ῌ QῦῦQ῍̮ῌ ῖQ ῍̮ῦ̰ῥQῌ Q̱ῳῤ̯̮Ῠῦ DNA/DNA ̲̱ ̲ ῼ̲QQῲῪῬῚ΅ῲῷῤ῏ΌῥΊῨΊῨῩ̯ ̮΅῍ ̯ῦ῭ῡ Ὼ῏ ῾ Ὸ̱̱̲ῲ ῏̲Ίῠ΅ ῷ̱ ῩQῦQQῴ ῏῾ῧῬ̯Ῑ'̯ 92 QQῦ̲῏ Ώ̮ῬQQῦQQῦQ῍ῧ 16S rDNA ̲Qῡ 98.7ΐ̳ ̮ῦ̲QῦQQΊ̯Q̮Ῥ῭Q̯ῠ΅ῢῦQQῲ῏ Ί̯῍ QQῦ̳ QQῧῗῦQῐῦῲῡQΰῡ̮΅ ̮ῌ QQQῌ Ὸῥ'ῺQQῡῧῗῦQQ̱ῧῥ ῦQQ̱ῥQ̮ῌ Q῍ῧQQῡ̮ῤῗῢ̮ῧ῍ QQ ῞̳QQῦῸQ̳QQῧ 2002 ̲ῥ ῞̳ῦQ῍ῦ ῲῥQ̱ῳῤ̯̮Ῠῦ DNA/DNA ̲̱̲ ῼ̲QQῥ Q΅Q῍ῦῲ̮QῢΊῠΎ῭ῧQῦ΅ housekeeping genes ῦῬ̯ῲQ̮̰̮̯ῢῦQQῲ῏Ίῌ Q QῳῥῧQ̳̱Qῦ̲῏ΏῦῬ̯̮Q̰̯ῠ΅῍ ῧQ ῌ501῍1195 Q̳QQQ 1῍1 ῼQ̮ ῞̳QQQQ Jpn.