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To thrive, organisms must maintain physiological and environmental variables in optimal ranges. However, in a dynamic world, the optimal range of a variable might fluctuate depending on the organism's state or environmental conditions. Given these fluctuations, how do biological control systems maintain optimal control of physiological and environmental variables? We explored this question by studying the phototactic behavior of larval zebrafish. We demonstrate, with behavioral experiments anddoi:10.1101/2020.07.18.210260 fatcat:xiigju3vcffajpawhcnela2vai