In systems accreting well below the Eddington rate, the plasma in the innermost regions of the disk is collisionless and two-temperature, with the ions hotter than the electrons. Yet, whether a collisionless faster-than-Coulomb energy transfer mechanism exists in two-temperature accretion flows is still an open question. We study the physics of electron heating during the growth of ion velocity-space instabilities, by means of multi-dimensional particle-in-cell (PIC) simulations. A large-scale

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... ompression - embedded in a novel form of the PIC equations - continuously amplifies the field. This constantly drives a pressure anisotropy P_perp > P_parallel, due to the adiabatic invariance of the particle magnetic moments. We find that, for ion plasma beta values beta_i ~ 5-30 appropriate for the midplane of low-luminosity accretion flows, mirror modes dominate if the electron-to-proton temperature ratio is > 0.2, whereas if it is < 0.2 the ion cyclotron instability triggers the growth of strong Alfven-like waves, that pitch-angle scatter the ions to maintain marginal stability. We develop an analytical model of electron heating during the growth of the ion cyclotron instability, which we validate with PIC simulations. We find that for cold electrons (beta_e < m_e/m_i), the electron energy gain is controlled by the magnitude of the E-cross-B velocity induced by the ion cyclotron waves. This term is independent of the initial electron temperature, so it provides a solid energy floor even for electrons starting with extremely low temperatures. On the other hand, the electron energy gain for beta_e > m_e/m_i - governed by the conservation of the magnetic moment in the growing fields of the instability - is proportional to the initial electron temperature. Our results have implications for two-temperature accretion flows as well as the solar wind and intracluster plasmas. [abridged]