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Pairing of food aroma with selected taste can lead to enhanced food flavor and eating euphoria, but how cross-modal sensory combinations are integrated to increase food reward value remains largely unclear. Here we report that combined stimulation by food aroma and taste drastically increased appetite in well-nourished larvae, and the appetizing effect involves a previously uncharacterized smell-taste integration process at axon terminals of two Gr43a gustatory neurons. Molecular geneticdoi:10.1101/859223 fatcat:ftmtnwdelfgijas4vsqsywycpe