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W. E. R. Rebholz, L. E. M. De Boer, M. Sasaki, R. H. R. Belterman, C. Nishida-Umehara. "The Chromosomal Phylogeny of Owls (Strigiformes) and New Karyotypes of Seven Species." CYTOLOGIA 58.4 (1993) 403-416