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Although cooperative behavior is generally acknowledged to occur among wolves, there is a lack of systematically collected data confirming the extent of development. lhe objectives of this study were to collect long-term, detailed observations documenting the role of social structure, seasonal influences, and individual participation in wolf pack cooperative activities. Individual cooperative strategies were associated with age, sex, and social positions and critically compared with results ofdoi:10.15760/etd.3213 fatcat:u5okwbjm2jf7tndesylkckpdoa