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The perianths of Liliaceae plants, such as lily and tulip, have two whorls of almost identical petaloid organs, which are called tepals. According to the modified ABC model proposed in tulip, the class B genes are expressed in whorl 1 as well as whorls 2 and 3, so that the organs of whorls 1 and 2 have the same petaloid structure. The floral structure of asparagus ( Asparagus officinalis L.) is similar to that of Liliaceae plants, however, the expression of B-class genes ( AODEF , AOGLOA ,doi:10.1266/ggs.80.95 pmid:16172521 fatcat:34iceywkcbhdvorbpoldp4wn7q