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Almost all higher organisms, including plants, insects, and mammals, are colonized by complex microbial communities and harbor a microbiome. Emerging studies with plants reveal that these microbiomes are structured and form complex, interconnected microbial networks. Within these networks, different taxa have different roles, and keystone species have been identified that could be crucial for plant health and ecosystem functioning. A new paper in this issue of PLOS Biology by Agler et al.doi:10.1371/journal.pbio.1002378 pmid:26871440 pmcid:PMC4752285 fatcat:be7yh5lyyvfrra5qw7vl3jjecm