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The thylakoid transmembrane ∆pH is the sole energy source driving translocation of precursor proteins by the ∆pH/Tat machinery. Consequently, proton translocation must be coupled to precursor translocation. For the precursor of the 17 kDa protein of the oxygen-evolving complex (pOE17), the protein translocation process is characterized by a steep drop in efficiency at an external pH below 7.0 and above 8.7. As the membrane ∆pH is virtually unaffected from pH 6.5 to 9.2, the loss in importdoi:10.1021/bi000115f pmid:10889030 fatcat:giondw5oujfrpj7sxy4ep6zlea