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<p><strong>Abstract.</strong> Morphological analysis of semen is a very important step in fertility assessment, but many semen defects are not detectable at the morphological level. These include pathological changes in sperm chromatin structure. During mammalian spermiogenesis, histone proteins associated with DNA structure are replaced by specific protamines, with which chromatin does not form nucleosomal complexes. In the fully developed, mature sperm, the histones are replaced withdoi:10.7482/0003-9438-57-013 fatcat:6yd7ll3mhbhodldrce72bqa2di