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Ascospore shape, wall layering, and surface structure are criteria considered important in separating Debaryomyces from certain other nitrate-negative yeast genera (Kreger-van Rij, 1970) . Transmission electron microscopy (TEM) showed ascospore walls of Debaryomyces to be composed of an outer thin, electron-dense layer and an inner layer that was much thicker, less electron-dense, and that formed the wartlike protrusions sometimes seen by light microscopy. In spores of D. hansenii (Zopf) Lodderdoi:10.2307/3759013 fatcat:o6wbe2hqxzbh3op7cjoonvq7o4