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In nature, division lines are delineated where multiple important environmental features change. These division lines may be singled out at the intersection of two geosystems (Balon 2000) where the functional uniformity of the geosystems located on both sides are preserved (Forman & Gordon 1986; Cadenasso et al. 2003). A significant environmental boundary is the upper forest boundary (timberline), which separates different vegetation zones: (1) forest from non-forest (Piękoś-Mirkowa & Mirekdoi:10.7163/gpol.0013 fatcat:xjdwa2v3tbcshoatjwi3khclgu