Eight pigeons were trained on a go-no go visual discrimination involving 1 S + and 15 S s. The 16 discriminative stimuli were black-and-white line drawings created by the factorial combination of 4 different geometric shapes (wedge, cylinder, cone, handle) in 4 different spatial locations (right, left, above, below) in relation to a common shape (cube). All of the pigeons readily learned this complex visual discrimination. Each bird's pecking behavior was controlled by both attributes of the

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... e drawings, but somewhat stronger stimulus control was exerted by the location of the added component than by its shape. Across all 8 pigeons, there was an inverse relation between stimulus control by component shape and component location. These results document pigeons' joint processing of "what" and "where" information in visual discrimination learning.