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KEYWORDS neurotropic virus, output network tracing, vesicular stomatitis virus, attenuated 2 cytotoxicity, enhanced anterograde trans-synaptic tracing 3 Abstract Understanding the connecting structure of brain network is the basis to reveal the principle of the brain function and elucidate the mechanism of brain diseases. Trans-synaptic tracing with neurotropic viruses has become one of the most effective technologies to dissect the neural circuits. Although the retrograde trans-synapticdoi:10.21203/rs.2.21708/v1 fatcat:s4ak3ex3yfbnlnzmsdzqwpwvoe