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RNA structural motifs can be identified using methods that analyze base–base interactions and the conformation of a structure's backbone; however, these approaches do not necessarily take into consideration the hydrogen bonds that connect the bases or the networks of inter-connected hydrogen-bonded bases that are found in RNA structures. Large clusters of RNA bases that are tightly inter-connected by a network of hydrogen bonds are expected to be stable and relatively rigid substructures. Suchdoi:10.3390/cryst9110550 fatcat:ov6m2rg5ordjje4otgwfwaq2fu