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In earth system modeling, biogeochemistry strongly affects mass and energy exchanges between ecosystems and the physical climate system (Heinze et al., 2019) . Morphologically, biogeochemistry has three pillars: biology, geophysics, and chemistry. In the context of mathematical modeling, geophysics and chemistry generally have much stronger theoretical foundations than biology (Brutsaert, 2005; Stumm & Morgan, 1996; Vallis, 2006) , even though all three are macroscale responses that emerge fromdoi:10.1029/2021ms002469 fatcat:b2n6kpog3fdatfxan75mq7mn3a