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The role of environmental variation in the evolution of lifespan is contested. Classic theory suggests that variable environments result in evolution of long life but novel theoretical breakthroughs show that environmental variation can instead select for short lifespan when the changes are rapid relative to generation time. Here we combined simulation modelling and experimental evolution to study the evolution of lifespan in response to temperature variation to provide the first empirical testdoi:10.1101/2020.01.29.924373 fatcat:lg27qokj7bcgxoszwnaol6n5xe