<p><strong>Abstract.</strong> Fire plays an important role in terrestrial ecosystems. The burning of biomass affects carbon and water fluxes and the distribution of vegetation. To understand the effect of the interactive processes of fire and ecological succession on land surface carbon and water fluxes, this study utilized the Community Land Model version 4.5 to conduct a series of experiments that included and excluded fire and dynamic vegetation processes. Results of the

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... eriments that excluded dynamic vegetation showed a global increase in net ecosystem production (NEP) in post-fire regions, which has been shown in previous studies with the similar modeling practices. However, inclusion of dynamic vegetation revealed a fire-induced decrease in NEP in some regions. Additionally, the carbon sink in post-fire regions reduced when the dominant vegetation type was changed from trees to grasses. This study shows that inclusion of dynamic vegetation enhances carbon emissions from fire by reducing terrestrial carbon sinks; however, this effect is somewhat mitigated by the increase in terrestrial carbon sinks when dynamic vegetation is not used. Results also show that fire-induced changes in vegetation modify the soil moisture profile because grasslands are more dominant in post-fire regions; this results in less moisture within top soil layers compared to non-burned regions, even though transpiration is reduced overall. These findings are different from those of previous fire model evaluations, that ignore vegetation dynamics, and thus highlight the importance of interactive processes between fire and vegetation dynamics, particularly when evaluating recent model developments with respect to fire and vegetation dynamics.</p>