Charles Darwin was not only the author of the theory of the evolution which basis is made by the concept of natural selection, but also the outstanding evolutionary ecologist. The contribution of Darwin to ecology is underestimated because in its works there is no conceptual framework of modern ecology. Natural selection and struggle for life in «Origin of species» of Ch. Darwin correspond the Volterra - Gause competitive exclusion principle. Darvinian «a place in the nature» to identically modern concept «ecological niche».