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Spatial environmental heterogeneity (EH) is considered one of the most important factors promoting species richness, but no general consent about the EH-richness relationship exists so far. This is because research methods and study settings vary widely, and because non-significant and negative associations have also been reported. My thesis provides a comprehensive review of the different measurements and terminologies of EH used in the literature, and presents strong quantitative evidence ofdoi:10.21425/f5fbg27952 fatcat:jzwcwlj6xjd23hhxvp77b6ixzy