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Genome-wide association studies (GWAS) allows to dissect the genetic basis of complex traits at the population level. However, despite the extensive number of trait-associated loci found, they often fail to explain a large part of the observed phenotypic variance. One potential source of this discrepancy could be the preponderance of undetected low-frequency genetic variants in natural populations. To increase the allele frequency of those variants and assess their phenotypic effects at thedoi:10.1101/609917 fatcat:pazbepaou5bcxbzjj6hyvdkqlu