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The spatial and temporal distribution of females is affected by the distribution of food and other resources such as nest sites, whereas spacing behavior of males is suggested to be determined by the distribution of females. When females are aggregated, the males should display territorial behavior in order to monopolize this defensible resource, while territorial, more dispersed females should lead males to have large, overlapping home ranges. However, previous studies found contrastingfatcat:otojuykycjdlpjutzhukpkzjve